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工作經(jīng)歷
2005.10 - 至今 ：華南農(nóng)業(yè)大學生命科學學院，教授
2004.08-2005.09：中國科學院華南植物園，研究員
1999.12-2004.07：中國科學院華南植物研究所，副研究員
1992.07-1999.11：中國科學院華南植物研究所，研究實習員，助理研究員
學習經(jīng)歷
博士 博士研究生 理學 植物學 香港大學 全日制 1996-2 ——1999-4
碩士 碩士研究生 理學 植物學 中國科學院華南植物園 全日制 1989-9 ——1992-4
學士 大學本科 理學 生物學 華中師范大學 全日制 1983-9 ——1987-6
研究興趣
植物系統(tǒng)發(fā)育與分子生態(tài)等方面的研究。
應用分子生物學數(shù)據(jù)，結(jié)合形態(tài)學，研究重要植物類群的親緣關(guān)系和性狀的演化，和特定生態(tài)條件下物種適應和分化的分子機制。
主持課題
1.國家自然科學基金項目：中國及泛喜馬拉雅地區(qū)紫金牛屬（報春花科）的分類學研究（項目批準號31570193）。2016.1-2019.12.
2.國家自然科學基金項目：點地梅屬（報春花科）的系統(tǒng)學與生物地理學（項目批準號31270260）。2013.1-2016.12.
3.國家自然科學基金項目：珍珠菜族（紫金牛科）的歷史生物地理學（項目批準號31070189）。2011.1-2013.12.
4.國家自然科學基金項目：報春花屬藏報春亞屬的界定和系統(tǒng)發(fā)育關(guān)系（項目批準號30870171）。2009.1-2011.12.
5.國家自然科學基金項目：基于分子系統(tǒng)發(fā)育推定冰期避難所和歷史遷移途徑：鄂報春、卵葉報春的親緣地理學（項目批準號40671066）。2007.1-2009.12.
6.國家自然科學基金項目：喜馬拉雅-橫斷山地區(qū)的物種形成：報春花屬脆蒴報春組的系統(tǒng)發(fā)育和親緣地理學（項目批準號30470125）。2005.1-2007.12.
7.廣東省自然科學基金項目：報春花屬脆蒴報春組的系統(tǒng)發(fā)育關(guān)系研究（項目批準號031255）。2004.1-2005.12.
8.國家自然科學基金項目：報春花屬脆蒴報春組的系統(tǒng)發(fā)育關(guān)系（項目批準號30370100）。2004.1-2004.12.
9.中國科學院知識創(chuàng)新工程重要方向性項目專題: 若干重要植物類群的系統(tǒng)發(fā)育重建和分子進化—五味子科系統(tǒng)發(fā)育關(guān)系的研究（項目批準號kscxz-sw-101A）。2002.1-2004.12.
10.國家自然科學基金項目：珍珠菜屬的分子系統(tǒng)學研究（項目批準號39970053）。2000.1-2002.12.
發(fā)表論文
1) Chen, S., X.K. Yan, G. Hao and Y. Xu*. 2019. The complete chloroplast genome of Primula tsiangii W. W. Smith (Primulaceae): a karst endemic primrose in Southwest China. Mitochondrial DNA Part B 4: 2627-2628, DOI: 10.1080/23802359.2019.1642160.
2) Xu, Y., G. Hao* and C.M. Hu. 2019. Primula vallicola sp. nov. (Primulaceae) from Sichuan, China. Nordic Journal of Botany. 2019: e02116.
3) Yan, X.K., H.F. Yan and G. Hao*. 2019. The complete chloroplast genome of Ardisia crenata Sims (Primulaceae): a popular ornamental plant in China. Mitochondrial DNA Part B, 4: 1790-1791, DOI: 10.1080/23802359.2019.1612292.
4) Liu, W.Z., H.H. Kong, J. Zhou, P.W. Fritsch, G. Hao* and W. Gong *. 2018. Complete chloroplast genome of Cercis chuniana (Fabaceae) with structural and genetic comparison to six species in Caesalpinioideae. International Journal of Molecular Sciences, 19: 1286, doi:10.3390/ijms19051286.
5) Wu, X., Y.S. Ye, H. Liu, C.M. Hu and G. Hao*. 2018. Ardisia medogensis (Primulaceae), a new species from Xizang, China. Phytotaxa 373: 91–94.
6) Xu, Y., G. Hao* and C.M. Hu. 2018. On the identity of Androsace sarmentosa var. yunnanensis (Primulaceae) from Yunnan, China. Nordic Journal of Botany. 20182018:e01886.
7) Huang, G.H., X.K. Yan, C.M. Hu and G. Hao*. 2018. Ardisia bullata (Primulaceae), a new species from Hainan, China. Annales Botanici Fennici 55: 329-332.
8) Yan, H.F., C.Y. Zhang, A.A. Anderberg, G. Hao, X.J. Ge* and J.J. Wiens*. 2018. What explains high plant richness in East Asia? Time and diversification in the tribe Lysimachieae (Primulaceae). New Phytologist 219: 436–448.
9) Huang, G.H., X.K. Yan and G. Hao*. 2018. Dichroa fistulosa (Hydrangeaceae), A New Species from Guangdong, China. Journal of Tropical and Subtropical Botany 26: 429-432.
10) Yuan, S., D.X. Zhang and G. Hao*. 2018. Primula chimingiana (Primulaceae), a new species from Sichuan, China. Nordic Journal of Botany. 2018: e01390.
11) Xu, Y., C.M. Hu and G. Hao*. 2017. Primula centellifolia (Primulaceae), a new species from south-central China. Phytotaxa 326: 259-263.
12) Huang, G.H., C.M. Hu and G. Hao*. 2017. Rediscovery of Ardisia gigantifolia and the reinstatement of A. kteniophylla (Primulaceae) – finding the correct name for the Chinese medicinal plant ’Zou Ma Tai’. Nordic Journal of Botany 35: 628-632.
13) Hu, C.M., G. Hao* and P. Van The. 2017. Ardisia phankelociana (Primulaceae), a new species from Vietnam. Annales Botanici Fennici 54: 361-363.
14) Xu, Y., G.H. Huang, C.M. Hu and G. Hao*. 2017. Primula pengzhouensis (Primulaceae), a new species from Sichuan, southwestern China. Plant Diversity 39: 229-231.
15) Huang, G.H., G. Hao* and C.M. Hu. 2017. Hymenandra A. DC. (Primulaceae), A New Generic Record for China. Journal of Tropical and Subtropical Botany 25: 282-284.
16) Huang, G.H. and G. Hao*. 2017. A new species of genus Stimpsonia (Primulaceae) from Guangdong, China. Nordic Journal of Botany 35: 157-161.
17) Yan, H.F., Y. Xu, Z.M. Zhu, C.M. Hu and G. Hao*. 2017. Lysimachia sinopilosa (Primulaceae), a new species from Yunnan, China. Annales Botanici Fennici 54: 45-48.
18) Zhang, C.Y., T.J. Liu, Y. Xu, H.F. Yan*, G. Hao and X.J. Ge. 2017. Characterization of the whole chloroplast genome of an endangered species Primula kwangtungensis (Primulaceae). Conservation Genet Resour 9: 87–89.
19) Liu, Y.J., C.Y. Zhang, G. Hao, X.J. Ge and H.F. Yan*. 2016. Characterization of novel microsatellite loci for Primula poissonii (Primulaceae) using high-throughput sequencing technology. Molecule 21(5), p.536.
20) Li, C.H., Y.J. Liu, C.Y. Zhang, H.F. Yan, X.J. Ge and G. Hao*. 2016. Characterization of Polymorphic Microsatellite Markers for Primula sikkimensis (Primulaceae) Using a 454 Sequencing Approach. Applications in Plant Sciences 4(7): 1600015.
21) Liu, T.J., C.Y. Zhang, H.F. Yan, X.J. Ge and G. Hao*. 2016. Complete plastid genome sequence of Primula sinensis (Primulaceae): structure comparison, sequence variation and evidence for accD transfer to nucleus. PeerJ 4:e2101; DOI 10.7717/peerj.2101.
22) Xu, Y., X.L. Yu, C.M. Hu and G. Hao*. 2016. Morphological and molecular phylogenetic data reveal a new species of Primula (Primulaceae) from Hunan, China. PLoS ONE 11(8): e0161172. DOI:10.1371/journal.pone.0161172.
23) Xu, Y., C.M. Hu and G. Hao*. 2016. Primula scopulicola (Primulaceae), a new species from Sichuan, southwestern China. Annales Botanici Fennici 53: 277-279.
24) Xu, Y., H.F. Yan, C.M. Hu and G. Hao*. 2016. Primula persimilis sp. nov. (Primulaceae) from Sichuan, China. Nordic Journal of Botany 34: 409-412.
25) Xu, Y., C.H. Li, C.M. Hu and G. Hao*. 2016. Primula wawushanica sp. nov. (Primulaceae) from Sichuan southwestern China. Nordic Journal of Botany 34: 156-158.
26) Xu, Y., Hu C.M. and G. Hao*. 2016. The pollen morphology of Androsace L. (Primulaceae) and its systematic implications. Journal of Systematics and Evolution 54: 48-64.
27) Xu, Y., T.J. Liu, C.M. Hu and G. Hao*. 2015. Primula anthemifolia (Primulaceae), a new species from Sichuan, China. Annales Botanici Fennici 52: 373-375.
28) Liu, Y.J., J. Liu, C.M. Hu and G. Hao*. 2015. Non-monophyly of Primula subgenera Auganthus and Carolinella (Primulaceae) as confirmed by the nuclear DNA sequence variation. Plant Systematics and Evolution 301: 2057-2071.
29) Yan, H.F., Y.J. Liu, X.F. Xie, C.Y. Zhang, C.M. Hu, G. Hao* and X.J. Ge*. 2015. DNA barcoding evaluation and its taxonomic implications in the species-rich genus Primula L. in China. PLoS ONE 10(4): e0122903. doi:10.1371/journal.pone.0122903.
30) Xu, Y., C.M. Hu and G. Hao*. 2015. A new species of Primula (Primulaceae) from Sichuan, China. Journal of Tropical and Subtropical Botany 23: 147-150.
31) Yu, X.L., Y. Xu, C.M. Hu and G. Hao*. 2015. Primula hunanensis sp. nov. (Primulaceae) from Hunan, south-central China. Nordic Journal of Botany 33: 381-384.
32) Xu, Y., S. Yuan, C.M. Hu and G. Hao*. 2014. Primula dejuniana (Primulaceae), a new species from Sichuan, China. Annales Botanici Fennici 51: 372-374.
33) Xu, Y., Z.Y. Liu, X.L. Yu, C.M. Hu and G. Hao*. 2014. Primula pelargoniifolia (Primulaceae), a new species from Chongqing, China. Annales Botanici Fennici 51: 125-127.
34) Xu, B. Q., G. Hao and N. H. Xia*. 2014. Saussurea haizishanensis (Compositae, Cardueae), a new species from Sichuan, China. Nordic Journal of Botany 32: 154-159.
35) Wu, X., Y. Xu, C.M. Hu and G. Hao*. 2013. Primula mianyangensis (Primulaceae), a new species from Sichuan, China. Phytotaxa 131: 49-52.
36) Xu, B.Q., N.H. Xia and G. Hao*. 2013. Saussurea bijiangensis (Asteraceae), a new species from Yunnan, China. Annales Botanici Fennici 50: 103-106.
37) Xu, B.Q., G. Hao and N.H. Xia*. 2013. Saussurea wenchengii (Compositae, Cardueae), a new species from Qinghai, China. Annales Botanici Fennici 50: 83-86.
38) Yan, H.F., C.Y. Zhang, F.Y. Wang, C.M. Hu, X.J. Ge and G. Hao*. 2012. Population Expanding with the Phalanx Model and Lineages Split by Environmental Heterogeneity: A Case Study of Primula obconica in Subtropical China. PLoS ONE 7(9): e41315. doi:10.1371/journal.pone.0041315. Epub 2012 Sept. 19.
39) Xie, X.F., H.F. Yan, F.Y. Wang, X.J. Ge, C.M. Hu and G. Hao*. 2012. Chloroplast DNA phylogeography of Primula ovalifolia in central and adjacent southwestern China: Past gradual expansion and geographical isolation. Journal of Systematics and Evolution 50: 284-294.
40) Yan, H.F. and G. Hao*. 2012. Lysimachia huchimingii sp. nov. (Primulaceae) from China. Nordic Journal of Botany 30: 443-445.
41) Zhang, C.Y., F.Y. Wang, H.F. Yan, G. Hao, C.M. Hu and X.J. Ge*. 2012. Testing DNA barcoding in closely-related groups of Lysimachia L. (Myrsinaceae). Molecular Ecology Resources 12: 98-108.
42) Hu, C.M. and G. Hao*. 2011. New and noteworthy species of Primula from China. Edinburgh Journal of Botany 68: 297-300.
43) Hu, C.M., G. Hao* and Y.F. Deng. 2011. Taxonomic and nomenclatural notes on the black-flowered Primula from China. Journal of Tropical and Subtropical Botany 19: 222-224.
44) Yan, H.F., G. Hao, C.M. Hu and X.J. Ge*. 2011. DNA barcoding in closely related species: a case study of Primula L. sect. Proliferae Pax (Primulaceae) in China. Journal of Systematics and Evolution 49: 225-236.
45) Mao, K.S., G. Hao, J.Q. Liu*, R.P. Adams and R.I. Milne. 2010. Diversification and biogeography of Juniperus (Cupressaceae): variable diversification rates and multiple intercontinental dispersals. New Phytologist 188: 254-272.
46) Li, L.F., M. H?kkinen, Y.M. Yuan, G. Hao and X.J. Ge. 2010*. Molecular phylogeny and systematics of the banana family (Musaceae) inferred from multiple nuclear and chloroplast DNA fragments, with a special reference to the genus Musa. Molecular Phylogenetics and Evolution 57: 1-10.
47) Yan, H.F., C.H. He, C.I. Peng, C.M. Hu and G. Hao*. 2010. Circumscription of Primula subgenus Auganthus (Primulaceae) based on chloroplast DNA sequences. Journal of Systematics and Evolution 48: 123-132.
48) Yan, H.F., X.J. Ge, C.M. Hu and G. Hao*. 2010. Isolation and characterization of microsatellite loci for ornamental plant, Primula obconica Hance (Primulaceae). HortScience 45: 314-315.
49) Huang, X.A.*, H.Y. Jiang and G. Hao. 2009. Direct HPLC detection of benzodilactones and quinones in glands of Lysimachia fordiana. Fitoterapia 80: 173-176.
50) Pan, Y.Z., L.Q. Fang, G. Hao, J. Cai and X. Gong*. 2009. Systematic positions of Lamiophlomis and Paraphlomis (Lamiaceae) based on nuclear and chloroplast sequences. Journal of Systematics and Evolution 47: 535-542.
51) 顏海飛，王鳳英，郝剛*. 2009. 報春花屬的保育遺傳學及親緣地理學研究進展. 廣西植物29: 751-757.
52) 寧淑萍，顏海飛，郝剛，葛學軍*. 2008. 植物DNA條形碼研究進展. 生物多樣性 16: 417-425.
53) Chen, G.Q., L.F. Li, G. Hao, S.H. Shi and X.J. Ge*. 2008. Characterization of seven genomic and one dbEST-derived microsatellite loci in the river mangrove Agceras corniculatum (Myrsinaceae). Conservation Genetics 9: 449-451.
54) Wang, F.Y., X.J. Ge, X. Gong, C.M. Hu and G. Hao*. 2008. Strong genetic differentiation of Primula sikkimensis in the East Himalaya-Hengduan Mountains. Biochemical Genetics 46: 75-87.
55) Wang, F.Y., X. Gong, C.M. Hu and G. Hao*. 2008. Phylogeography of an alpine species Primula secundiflora inferred from the chloroplast DNA sequence variation. Journal of Systematics and Evolution 46: 13-22.
56) Yan, H.F., C.I. Peng, C.M. Hu and G. Hao*. 2007. Phylogeographic structure of Primula obconica (Primulaceae) inferred from chloroplast microsatellites (cpSSRs) markers. Acta Phytotaxonomica Sinica 45: 488-496.
57) Wang, X.L., T.Y. Chiang, N. Roux, G. Hao and X.J. Ge*. 2007. Genetic diversity of wild banana (Musa balbisiana Colla) in China as revealed by AFLP markers. Genetic resources and Crop Evolution 54: 1125-1132.
58) Liu, Q., P.M. Peterson, J.T. Columbus, N.X. Zhao*, G. Hao and D.X. Zhang. 2007. Inflorescence diversification in the finger millet clade (Chloridoideae, Poaceae): a comparison of molecular phylogeny and developmental morphology. American Journal of Botany 94: 1230-1247.
59) 張巧玲，謝建光，陳顯化，郝剛*. 2007. 中國紫金牛屬圓齒組果實微形態(tài)特征研究. 武漢植物研究 25: 432-439.
60) 許炳強，夏念和，王少平，郝剛*. 2007. 中國木犀屬植物葉脈形態(tài)及其分類學意義. 廣西植物27: 697-705.
61) 張巧玲，王少平，胡啟明，郝剛*. 2007. 中國紫金牛屬圓齒組花粉形態(tài)研究及其分類學意義. 廣西植物 27: 385-392.
62) 許炳強，夏念和，王少平，郝剛*. 2007. 中國木犀屬植物葉表皮形態(tài). 武漢植物研究25: 1-10.
63) Liu, Z., G. Hao, Y.B. Luo, L.B. Thien, S.W. Rosso, A.M. Lu and Z.D. Chen. 2006. Phylogeny and androecial evolution in Schisandraceae, inferred from sequences of nuclear ribosomal DNA ITS and chloroplast DNA trnL-F regions. International Journal of Plant Sciences 167: 539-550.
64) Hu, C.M. G. Hao*. 2006. A new synonym of Primula boreiocalliantha Balf. f. & Forrest (Primulaceae). Acta Phytotaxonomica Sinica 44: 477-479.
65) Han, R.L., D.X. Zhang* and G. Hao. 2006. Two new species of Viscum (Viscaceae) from China. Nordic Journal of Botany 23: 719-724.
66) Hu, C.M., G. Hao* and Y.F. Deng. 2006. The identity of Lysimachia longshengensis G. Z. Li & S. C. Tang. Acta Phytotaxonomica Sinica 44: 195-196.
67) 劉青，趙南先*，郝剛. 2006. 禾本科虎尾草亞科細胞學研究分析. 熱帶亞熱帶植物學報 14: 347-353.
68) Hu, C.M*. and G. Hao. 2005. Notes on the genus Sabia (Sabiaceae) from Vietnam. Blumea 50: 551-553.
69) Xiao, L.Q., X. Gong*, G. Hao, X.J. Ge, B. Tian and S.X. Zheng. 2005. Comparison of genetic diversity in two species of Cycas. Australian Journal of Botany 59: 219-223.
70) Liu, Q., N.X. Zhao*, G. Hao, X.Y. Hu and Y.X. Liu. 2005. Caryopsis morphology of the Chloridoideae (Gramineae) and its systematic implications. Botanical Journal of Linnean Society 148: 57-72.
71) Liu, Q., N.X. Zhao* and G. Hao. 2005. Inflorescence structures and evolution in subfamily Chloridoideae. Plant Systematics and Evolution 251: 183-198.
72) Ge, X. J., L.B. Zhang, Y.M. Yuan, G. Hao and T.Y. Chiang*. 2005. Strong genetic differentiation of the East-Himalayan Megacodon stylophorus (Gentianaceae) detected by Inter-Simple Sequence Repeats (ISSR). Biodiversity and Conservation 14: 849-861.
73) Liu, Q., N.X. Zhao* and G. Hao. 2005. Pollen morphology of Eustachys tenera (Chloridoideae, Gramineae). Pakistan Journal of Botany 37: 503-506.
74) 顏海飛，王小蘭，胡啟明，郝剛*。2005. 鄂報春（Primula obconica Hance）親緣地理學的初步研究. 熱帶亞熱帶植物學報13: 526-532.
75) 許炳強，鄧云飛，郝剛*. 2005。厚葉木犀名稱的合格發(fā)表. 熱帶亞熱帶植物學報13: 451.
76) 王鳳英，葛學軍，郝剛*，胡啟明. 2005. 喜馬拉雅- 橫斷山區(qū)鐘花報春居群遺傳多樣性及遺傳分化. 熱帶亞熱帶植物學報 13: 149-153.
77) 許炳強，郝剛*，胡曉穎. 2005. 中國木犀屬花粉形態(tài)研究及其系統(tǒng)學意義. 熱帶亞熱帶植物學報13: 29-39.
78) 劉青，趙南先*，郝剛. 2005. 虎尾草亞科系統(tǒng)發(fā)育關(guān)系的初步研究. 熱帶亞熱帶植物學報13: 432-442.
79) Hao, G., Y.M. Yuan*, C.M. Hu, X.J. Ge and N.X. Zhao. 2004. Molecular Phylogeny of Lysimachia (Myrsinaceae s.l.) based on chloroplast trnL-F and nuclear ribosomal ITS sequences. Molecular Phylogenetics and Evolution 31: 323-339.
80) Liu, Q., N.X. Zhao* and G. Hao. 2004. Pollen morphology of the Chloridoideae (Gramineae). Grana 43: 1-10.
81) Xue, D.W., X.J. Ge, G. Hao and C.Q. Zhang*. 2004. High genetic diversity in a rare, narrowly endemic primrose species: Primula interjacens (Primulaceae). Acta Botanica Sinica 46: 1163-1169.
82) Xiao, L.Q., X.J. Ge, X. Gong*, G. Hao and S.X. Zhang. 2004. ISSR variation in the endemic and endangered plant Cycas guizhouensis (Cycadaceae). Annals of Botany 94: 133-138.
83) Wang D.L., Z.C. Li, G. Hao and X.J. Ge*. 2004. Genetic diversity of Calocedrus macrolepis (Cupressaceae) in southwestern China. Biochemical Systematics and Ecology 32: 797-807.
84) 王玉金，李小娟，郝剛，劉建全*. 2004. 點地梅屬的分子系統(tǒng)學、生物地理學和墊狀形態(tài)的趨同進化. 植物分類學報 42: 481-499.
85) 韓容蘭，張奠湘*，郝剛. 2004. 桑寄生科花粉形態(tài)的研究. 植物分類學報 42: 436-456.
86) Han, R.L., D.X. Zhang and G. Hao. 2004. Interfamilial relationships of Santalales as revealed by chloroplast trnL intron sequences. Journal of Tropical and Subtropical Botany 12: 393-398.
87) Hao, G., D.X. Zhang*, M.Y. Zhang, L.X. Guo and S.J. Li. 2003. Phylogenetics of Bauhinia subgenus Phanera (Leguminosae: Caesalpinioideae) based on ITS sequences of nuclear ribosomal DNA. Botanical Bulletin of Academia Sinica 44: 223-228.
88) 韓榮蘭，張奠湘*，郝剛. 2003. 中國槲寄生屬花粉形態(tài)的研究. 廣西植物 23: 318-320.
89) 韓榮蘭，張奠湘*，郝剛. 2002. 中國槲寄生屬植物及其寄主的地理分布. 熱帶亞熱帶植物學報 10: 222-228.
90)Hao, G., C.M. Hu* and Nam Sook Lee. 2002. Circumscriptions and phylogenetic relationships of Primula sect. Auganthus and Ranunculoides: evidence from nrDNA ITS sequences. Acta Botanica Sinica 44:72-75.
91) Hao, G., Dong-Hee Lee, June Seung Lee and Nam Sook Lee*. 2002. A study of taxonomical relationships among species of Korean Allium sect. Sacculiferum (Alliaceae) and related species using inter-simple sequence repeat (ISSR) markers. Botanical Bulletin of Academia Sinica 43: 63-68.
92) Hao, G., D. X. Zhang*, L.X. Guo, M.Y. Zhang, Y.F. Deng and X.Y. Wen. 2001. A phylogenetic and biogeographic study of Cercis (Leguminosae). Acta Botanica Sinica 43:1275-1278.
93) 郝剛，胡啟明*. 2001. 珍珠菜屬系統(tǒng)發(fā)育關(guān)系的初步研究. 熱帶亞熱帶植物學報 9: 93-100.
94) Hao, G., M.L. Chye and R.M.K. Saunders*. 2001. A phylogenetic analysis of the Schisandraceae based on morphology and nuclear ribosomal ITS sequences. Botanical Journal of Linnean Society 135: 401-411.
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